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Family animation. New York stowaways Alex the lion, Marty the zebra, Melman the giraffe and Gloria the hippo are still trying to wing their way back to the Big Apple from Madagascar. The quartet reach Monte Carlo but attract the attention of an animal control officer. In their efforts to escape, the gang bundle onto a circus train and convince the circus animals they are Big Top entertainers, all part of a crazy plan that will have them clowning and high-wiring all the way back to America.

Featuring the voices of David Schwimmer (Melman), Jada Pinkett Smith (Gloria), Sacha Baron Cohen (Julien), Cedric the Entertainer (Maurice), Andy Richter (Mort), Tom McGrath (Skipper/First Policeman), Frances McDormand (Captain Chantel DuBois), Jessica Chastain (Gia), Bryan Cranston (Vitaly), Martin Short (Stefano), Chris Miller (Kowalski) and Christopher Knights (Private).