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Boris Johnson has made a "full and unreserved" apology to MPs for failing to declare more than £52,000 in income.

The former foreign secretary was ordered to apologise by the Committee on Standards over late declaration of book royalty payments.

He made a brief statement to the Commons ahead of the latest debate on Theresa May's EU deal.

Speaker John Bercow thanked him for his "prompt" response and said that was the end of the matter.

Parliament's Standards Commissioner Kathryn Stone had earlier ruled that the breaches were "neither inadvertent nor minor".

She referred the case to the Committee on Standards to decide on a punishment for Mr Johnson.

MPs are required to declare payments for any work carried out in addition to their duties as an MP, in the Register of Members' Interests.

Ms Stone found that Mr Johnson's registrations were late on four separate occasions, involving nine payments, "which suggested a lack of attention to, or regard for, the House's requirements".

"The committee concluded that Mr Johnson breached the rules of the House by failing to register remuneration within the required timetable on nine occasions,"

"In considering the appropriate sanction, it took into account what it considered to be aggravating and mitigating factors, and recommended that Mr Johnson make an apology to the House on a point of order."

The late declarations included a series of rights and royalty payments from Bulgaria, Hungary, the Czech Republic, France and the US for books "already written" by Mr Johnson.

The committee said aggravating factors in calling for Mr Johnson to apologise included the size of the sums involved and the number of breaches.

As a long-standing MP who had been a senior minister, he "could be expected to set an example within the House", it added.

Mr Johnson responded "promptly and helpfully" when the issue was raised with him, apologised to the Commissioner and put in place "effective measures to ensure that no further breach occurs", the committee added.

Making his apology, Mr Johnson told MPs the payments were "mainly unexpected foreign royalties, which I am very sorry to say were recorded late on the register of Members' interests".

He added that he was "grateful to the committee for recognising that there was no intention to mislead the House and that I had been completely transparent".